Cetacea
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{{Taxobox
| color = pink
| name = Cetaceans
| image = humpbackwhale37.jpg
| image_width = 250px
| image_caption = Humpback Whale breaching
| regnum = Animalia
| phylum = Chordata
| classis = Mammalia
| ordo = Cetacea
| ordo_authority = Brisson, 1762
| subdivision_ranks = Suborders
| subdivision =
Mysticeti
Odontoceti
(see text for families)
}}
Template:Portal
The order Cetacea includes whales, dolphins and porpoises.
Cetus is Latin and is used in biological names to mean "whale"; its original meaning, "large sea animal," was more general. It comes from Greek ketos ("sea monster"). Cetology is the branch of marine science associated with the study of cetaceans.
Cetaceans are the mammals most fully adapted to aquatic life. Their body is fusiform (spindle-shaped). The forelimbs are modified into flippers. The tiny hindlimbs are vestigial; they do not attach to the backbone and are hidden within the body. The tail has horizontal flukes. Cetaceans are nearly hairless, and are insulated by a thick layer of blubber.
The order Cetacea contains eighty-one known species, divided into two suborders, Mysticeti (baleen whales) and Odontoceti (toothed whales, which includes dolphins and porpoises).
Contents |
Evolution
Cetaceans (see-taý she-ańs)(L. cetus, whale) evolved from land mammals (most likely from certain hoofed carnivores which also gave rise to the artiodactyls - the even-hoofed mammals, including pigs and the hippopotamus) that once again adapted to marine life about 50 million years ago.
Adaptations for sea life
Over a period of millions of years, the cetacean returned to the sea - there was more food there, and more space than on land. Because of this increase in space, there was no natural limit to the cetacean's size (i.e. the amount of weight its legs could hold) since the water provided buoyancy. It had no longer any need for legs.
During this time, the cetacean lost the qualities that fitted it for land existence and gained new qualities for life at sea. Its hind limbs disappeared, its body became more tapered and streamlined - a form that enabled it to move swiftly through the water. The cetacean's original tail was replaced by a pair of flukes that acted like a propeller.
As part of this streamlining process, the bones in the cetacean's front limbs fused together. In time, what had been the forelegs became a solid mass of bone, blubber and tissue, making very effective flippers that balance the cetacean's tremendous bulk.
To preserving body heat in cold oceanic waters, the cetacean developed blubber, a thick layer of fat between the skin and the flesh that also acts as an emergency source of energy. In some cetaceans the layer of blubber can be more than a foot thick. No longer needed for warmth, the cetacean's fur coat disappeared, further reducing the resistance of the giant body to the water.
The ear bone called the hammer (malleus) is fused to the walls of the bone cavity where the ear bones are, making hearing in air as good as impossible. Instead sound are transmitted through their jaws and skull bones.
Breathing, seeing, hearing and echolocation
Since the cetacean is a mammal, it needs air to breathe. Because of this, it needs to come to the water's surface to exhale its carbon dioxide and inhale a fresh supply of oxygen. Naturally it cannot breathe under water, so as it dives a muscular action closes the blowholes (nostrils), which remain closed until the cetacean next breaks the surface. When it does, the muscles open the blowholes and warm air is exhaled.
To make this easier, the cetacean's blowholes have moved to the top of its head, giving it a quicker chance to expel the stale air and inhale fresh air. When the stale air, warmed from the lungs, is exhaled it condenses and vapourises as it meets the cold air outside. This is rather like when you breathe out on a cold day and a small cloud of warm air appears. This is called the 'blow', or 'spout', and each cetacean's blow is different in terms of shape, angle and height. This is how cetaceans can be identified at a distance by experienced whalers or whale-watchers.
The cetacean's eyes are set well back and to either side of its huge head. This means that cetaceans with pointed 'beaks' (such as dolphins) have good binocular vision forward and downward, but others with blunt heads (such as the Sperm Whale) can see either side but not directly ahead or directly behind. The eyes shed greasy tears which protect them from the salt in the water. Cetaceans also have an almost spherical lens in their eyes, which is most efficient at focusing what little light there is. However, their vision is generally quite poor (with the exception of the dolphin), which cetaceans make up with their excellent hearing abilities
Akin to the eyes, the cetacean's ears are also small. Life in the sea accounts for the cetacean's loss of its external ears, whose function is to collect sound waves and focus them in order for them to become strong enough to hear well. However, sound waves travel faster through the water than in the air, and so the external ear was no longer needed, and is no more than a tiny hole in the skin, just behind the eye. The inner ear, however, has become so well developed that the cetacean can not only hear sounds tens of miles away, but it can also discern from which direction the sound comes.
Some cetaceans are capable of echolocation. Mysticeti have little need of it, as they prey upon small fish that would be impractical to locate with echolocation. Some members of Odontoceti, such as dolphins and porpoises, perform echolocation. Many toothed whales emit clicks similar to those in echolocation, but we have yet to prove that they echolocate. Cetaceans use sound in the same way as bats - they emit a sound (called a click), which then bounces off an object and returns to them. From this, cetaceans can discern the size, shape, surface characteristics and movement of the object, as well as how far away it is. With this ability cetaceans can search for, chase and catch fast-swimming prey in total darkness. It is so advanced that most cetaceans can discern between prey and non-prey (such as humans or boats), and captive cetaceans can be trained to distinguish between, for example, balls of different sizes or shapes.
Cetaceans also use sound to communicate, whether it be groans, moans, whistles, clicks or the complex 'singing' of the Humpback Whale that is becoming so popular on wildlife documentaries and relaxation tapes.
Feeding
When it comes to food and feeding, cetaceans can be separated into two distinct groups. The 'toothed whales' or Odontoceti usually have lots of teeth that they use for catching fish, squid or other marine life. They do not chew their food, but swallow it whole. In the rare cases that they catch large prey, such as Killer Whales (Orcinus orca) catching a fur seal, they tear 'chunks' of it that in turn are swallowed whole. Some examples of the cetaceans in this group include sperm whales, beluga whales, dolphins and porpoises.
The 'baleen whales' or Mysticeti do not have teeth. Instead they have plates made of keratin (the same substance as our fingernails) which hang down from the upper jaw. These plates act like a giant filter, straining small animals (such as krill and fish) from the seawater. Cetaceans included in this group include the Blue Whale, the Humpback Whale, the Bowhead Whale and the Minke Whale.
It is a fallacy to believe that all Mysticeti feed on plankton, because for some species their prey animals are not plankton. The larger whales tend to eat small shoaling fish, such as herrings and sardine. These fish are called micronecton, and not plankton. One species of Mysticeti, the Gray Whale (Escrichtius robustus), is a benthic feeder, primarily eating sea floor crustaceans.
Misconceptions
A common misconception about cetaceans is that they are fish. That is incorrect— rather than being fish, they are in fact mammals.
As mammals, cetaceans have these characteristics that are common to all mammals:
- They are warm-blooded animals.
- They breathe in air through their lungs.
- They bear their young alive (other than monotremes)...
- ...and suckle them on their own milk.
- They have hair - though generally only a few 'whiskers'.
Another way of discerning a cetacean from a fish is by the shape of the tail. The tail of a fish is vertical and moves from side to side when the fish swims. The tail of a cetacean - called a "fluke" - is horizontal and moves up and down, as cetaceans' spines bend in the same manner as a human spine.
Taxonomic listing
- ORDER CETACEA
- Suborder Mysticeti: Baleen whales
- Family Balaenidae: Right whales and Bowhead Whale
- Genus Balaena
- Bowhead Whale, Balaena mysticetus
- Genus Eubalaena
- Atlantic Northern Right Whale, Eubalaena glacialis
- Pacific Northern Right Whale, Eubalaena japonica
- Southern Right Whale, Eubalaena australis
- Genus Balaena
- Family Balaenopteridae: Rorquals
- Subfamily Balaenopterinae
- Genus Balaenoptera
- Northern Fin Whale, Balaenoptera physalus
- Southern Fin Whale, Balaenoptera quoyi
- Northern Sei Whale, Balaenoptera borealis
- Southern Sei Whale, Balaenoptera schlegllii
- Bryde's Whale, Balaenoptera brydei
- Eden's Whale Balaenoptera edeni
- Northern Blue Whale, Balaenoptera musculus
- Southern Blue Whale, Balenoptera intermedia
- Indian Blue Whale, Balaenoptera indica
- Pygmy Blue Whale, Balaenoptera brevicauda
- Northern Minke Whale, Balaenoptera acutorostrata
- Southern Minke Whale, Balaenoptera bonaerensis
- Dwarf Minke Whale, Balaenoptera sp.
- Omura's Whale, Balaenoptera omurai
- Genus Balaenoptera
- Subfamily Megapterinae
- Genus Megaptera
- Humpback Whale, Megaptera novaeangliae
- Genus Megaptera
- Subfamily Balaenopterinae
- † Genus Eobalaenoptera
- † Harrison's Whale, Eobalaenoptera harrisoni
- Family Eschrichtiidae
- Genus Eschrichtius
- Gray Whale, Eschrichtius robustus
- Genus Eschrichtius
- Family Neobalaenidae: Pygmy Right Whale
- Genus Caperea
- Pygmy Right Whale, Caperea marginata
- Genus Caperea
- Family Balaenidae: Right whales and Bowhead Whale
- Suborder Odontoceti: toothed whales
- Superfamily Platanistoidea: River dolphins
- Family Iniidae
- Genus Inia
- Amazon River Dolphin, Inia geoffrensis
- Orinoco River Dolphin, Inia humboldtiana
- Bolivian River Dolphin, Inia boliviensis
- Genus Inia
- Family Lipotidae
- Genus Lipotes
- Chinese River Dolphin, Lipotes vexillifer
- Genus Lipotes
- Family Platanistidae
- Genus Platanista
- Ganges and Indus River Dolphin, Platanista gangetica
- Genus Platanista
- Family Pontoporiidae
- Genus Pontoporia
- La Plata Dolphin, Pontoporia blainvillei
- Pygmy La Plata Dolphin, Pontoporia tenuirostris
- Genus Pontoporia
- Family Iniidae
- Family Monodontidae
- Genus Monodon
- Narwhal, Monodon monoceros
- Genus Delphinapterus
- Beluga, Delphinapterus leucas
- Genus Monodon
- Family Phocoenidae: Porpoises
- Genus Neophocaena
- Black Finless Porpoise, Neophocaena phocaenoides
- White Finless Porpoise, Neophocaena asiaorientalis
- Genus Phocoena
- Harbour Porpoise, Phocoena phocaena
- Vaquita, Phocoena sinus
- Spectacled Porpoise, Phocoena dioptrica
- Burmeister's Porpoise, Phocoena spinipinnis
- Genus Phocoenoides
- Dall's Porpoise, Phocoenoides dalli
- Genus Neophocaena
- Family Physeteridae: Sperm Whale family
- Genus Physeter
- Sperm Whale, Physeter macrocephalus
- Genus Physeter
- Family Kogiidae
- Genus Kogia
- Dwarf Sperm Whale, Kogia sima
- Pygmy Sperm Whale, Kogia breviceps
- Genus Kogia
- Family Ziphidae, Beaked whales
- Genus Ziphius
- Cuvier's Beaked Whale, Ziphius cavirostris
- Genus Berardius
- Arnoux's Beaked Whale, Berardius arnuxii
- Baird's Beaked Whale (North Pacific Bottlenose Whale), Berardius bairdii
- Genus Tasmacetus
- Tasman Beaked Whale (Shepherd's Beaked Whale), Tasmacetus shepherdi
- Subfamily Hyperoodontidae
- Genus Indopacetus
- Indo-Pacific Beaked Whale (Longman's Beaked Whale), Indopacetus pacificus
- Genus Hyperoodon
- Northern Bottlenose Whale, Hyperoodon ampullatus
- Southern Bottlenose Whale, Hyperoodon planifrons
- Genus Mesoplodon, Mesoplodont Whale
- Hector's Beaked Whale, Mesoplodon hectori
- True's Beaked Whale, Mesoplodon mirus
- Gervais' Beaked Whale, Mesoplodon europaeus
- Sowerby's Beaked Whale, Mesoplodon bidens
- Gray's Beaked Whale, Mesoplodon grayi
- Pygmy Beaked Whale, Mesoplodon peruvianus
- Andrews' Beaked Whale, Mesoplodon bowdoini
- Spade Toothed Whale, Mesoplodon traversii
- Hubbs' Beaked Whale, Mesoplodon carlhubbsi
- Ginkgo-toothed Beaked Whale, Mesoplodon ginkgodens
- Stejneger's Beaked Whale, Mesoplodon stejnegeri
- Layard's Beaked Whale, Mesoplodon layardii
- Blainville's Beaked Whale, Mesoplodon densirostris
- Perrin's Beaked Whale, Mesoplodon perrini
- Genus Indopacetus
- Genus Ziphius
- Family Delphinidae: Dolphin
- Genus Cephalorhynchus
- Commerson's Dolphin, Cephalorhyncus commersonii
- Chilean Dolphin, Cephalorhyncus eutropia
- Heaviside's Dolphin, Cephalorhyncus heavisidii
- Hector's Dolphin, Cephalorhyncus hectori
- Genus Steno
- Rough-toothed Dolphin, Steno bredanensis
- Genus Sousa
- Atlantic Humpback Dolphin, Sousa teuszi
- Indo-Pacific Humpback Dolphin, Sousa chinensis
- Genus Sotalia
- River Tucuxi, Sotalia fluviatilis
- Marine Tucuxi, Sotalia guianensis
- Genus Tursiops
- Common Bottlenose Dolphin, Tursiops truncatus
- Indian Ocean Bottlenose Dolphin, Tursiops aduncus
- South American Bottlenose Dolphin, Tursiops gephyreus
- Genus Stenella
- Pantropical Spotted Dolphin, Stenella attenuata
- Atlantic Spotted Dolphin, Stenella frontalis
- Spinner Dolphin, Stenella longirostris
- Clymene Dolphin, Stenella clymene
- Striped Dolphin, Stenella coeruleoalba
- Genus Delphinus
- Short-beaked Common Dolphin, Delphinus delphis
- Long-beaked Common Dolphin, Delphinus capensis
- Genus Lagenodelphis
- Fraser's Dolphin, Lagenodelphis hosei
- Genus Lagenorhynchus
- White-beaked Dolphin, Lagenorhynchus albirostris
- Atlantic White-sided Dolphin, Lagenorhynchus acutus
- Pacific White-sided Dolphin, Lagenorhynchus obliquidens
- Dusky Dolphin, Lagenorhynchus obscurus
- Black-chinned Dolphin, Lagenorhynchus australis
- Cook Islands Dolphin, Lagenorhynchus sp.
- Hourglass Dolphin, Lagenorhynchus cruciger
- Genus Lissodelphis
- Northern Right Whale Dolphin, Lissodelphis borealis
- Southern Right Whale Dolphin, Lissodelphis peronii
- Genus Grampus
- Risso's Dolphin, Grampus griseus
- Genus Peponocephala
- Melon-headed Whale, Peponocephala electra
- Genus Feresa
- Pygmy Killer Whale, Feresa attenuata
- Genus Pseudorca
- False Killer Whale, Pseudorca crassidens
- Genus Orcinus
- Orca, Orcinus orca
- Genus Globicephala
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhyncus
- Genus Orcaella
- Irrawaddy Dolphin, Orcaella brevirostris
- Australian Snubfin Dolphin, Orcaella heinsohni
- Genus Cephalorhynchus
- Superfamily Platanistoidea: River dolphins
- Suborder Mysticeti: Baleen whales
References
- Rice, Dale W. (1998). Marine mammals of the world: systematics and distribution. Society of Marine Mammalogy Special Publication Number 4. 231 pp. See the Society's website for further details.
External links
Template:Wikispecies Template:Wikibookspar
- ARKive - images and movies of Whales, Dolphins and Porpoises
- American Cetacean Society
- British Cetacean Site especially interesting is taxonomy
- Cetacea.org homepage
- Walker's Mammals of the World Online - Cetaceans
- Tursiops.org: Current Cetacean-related news
- Whales, Dolphins, Porpoises and Cetaceans Forum
Mammals |
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Monotremata |
Marsupialia: Didelphimorphia | Paucituberculata | Microbiotheria | Notoryctemorphia | Dasyuromorphia | Peramelemorphia | Diprotodontia |
Placentalia: Afrosoricida | Macroscelidea | Tubulidentata | Hyracoidea | Proboscidea | Sirenia | Cingulata | Pilosa | Scandentia | Dermoptera | Primates | Rodentia | Lagomorpha | Insectivora | Chiroptera | Pholidota | Carnivora | Perissodactyla | Artiodactyla | Cetacea |
ar:حوت bg:Китоподобни ca:Cetaci cs:Kytovci cy:Morfil da:Hvaler de:Wale es:Cetáceo eo:Cetacoj fa:آببازان fr:Cetacea ko:고래 hr:Kitovi it:Cetacea he:לווייתנאים la:Cetacea lt:Banginis li:Walvèsechtege ms:Ikan paus nl:Walvissen ja:クジラ目 pl:Walenie pt:Cetáceos ru:Китообразные scn:Citacea sk:Veľryby (rad) sr:Плави кит fi:Valas sv:Valar zh:鲸